Hunters have killed two radio-collared wolves that roamed Grand Teton National Park, localizing a debate about the legal killing of "park" wolves used for research.

According to a story published by the Associated Press Wednesday, wildlife officials say it's unavoidable that park wolves will range outside park boundaries. Wyoming Game and Fish Department harvest data show 13 wolves reported killed in hunt areas bordering the park.

On Oct. 1, the Fish and Wildlife Service ended Endangered Species Act protection for Wyoming wolves.

Wildlife managers and pro-wolf groups are at odds following the shooting of at least 10 collared animals that frequent Wyoming's two national parks.

Eight collared animals have been shot outside of Yellowstone National Park. On Monday, Montana closed a portion of its hunt area abutting the park.